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Pertica Quadrifaria: Maine's State Fossil

State Capital: Augusta
State Nickname: Pine Tree State
State Motto: Dirigo ("I lead")
State Bird: Chickadee
State Floral: White pine cone & tassel
State Fossil: Pertica quadrifaria
State Cat: Maine Coon Cat
State Fish: Landlocked Salmon
State Insect: Honeybee
State Tree: White Pine
State Animal: Moose
State Gemstone: Tourmaline

Pertica quadrifaria is the scientific name of a primitive plant that lived about 390 million years ago during the Devonian Period. Its fossilized remains were discovered in 1968 in the rocks of the Trout Valley Formation in Baxter State Park near Mount Katahdin. Based on the type of rock it is found in today and the other fossils associated with it, Pertica quadrifaria grew in a brackish marsh near an active volcano. Fragments of the plants were preserved when they fell into the marsh and were covered by sediment before they could decay. After millions of years of burial, the plant remains are now exposed along eroding stream banks. The photo shows the central stem and several branches which terminate in sporangia (click on the photo for an enlarged view).

The plant illustrated at right is an artist's reconstruction of Pertica quadrifaria. It probably reached a maximum height of about six feet, making it the largest land plant at that time (Pertica is a Latin word meaning a "long pole or rod"). Its stem, which measured up to one inch in diameter, bore both sterile and fertile branches arranged in four rows which spiraled up the stem (quadrifaria means "in four ranks"). The fertile branches ended in dense clusters of sporangia, or spore cases, while the sterile branches subdivided to form forked tips. These forked ends may represent the first step in the evolution of leaves.

Pertica quadrifaria was selected as the Maine State Fossil for several reasons. It was first discovered in Maine. It is also a rare fossil; well-preserved remains of Pertica are found at only three other places in the world besides Maine. Finally, it is an important fossil scientifically as it represents a significant early step in the evolution of vascular land plants which ultimately resulted in numerous modern species, including the pine tree.

content courtesy of maine.gov